Wither

Progression

 * 1) Duration is measured in seconds.

Acquisition
Signet of Capture:
 * Desnas Hubor (Tangle Root only during Defend Denravi)
 * Evirso Sectus (Mineral Springs only during Defend Droknar's Forge)
 * Ignis Effigia (Perdition Rock)
 * Ludo Ossidi (Dragon's Gullet only during The Titan Source)